The four-year Graduate Programme is designed to train future group leaders in molecular cell biology research or their alternative chosen pathway. The LMCB provides an outstanding scientific environment and cutting edge technology. The LMCB is one of the four departments of the faculty of Life Sciences at UCL and it is strategically located at the heart of London main scientific institutions. The LMCB graduate programme, established in 1996, now includes about 30 investigators who participate in the training of Ph.D. candidates. The goals of our training are to produce scientists who have explored one area and one level of analysis in great depth, but who are familiar with the full scope of molecular and cell biology. They should be able to move from one level to another in a critical and creative manner. We also try to develop an appreciation for translational research that bears on human disease. Graduate students spend four years in the Programme and are entirely supported by an MRC grant. During their first year, students have a free choice of three laboratories from the groups within the LMCB plus additional participating labs from other departments in UCL, and spend three months in each, working on research projects. They also participate in twice weekly tutorials that are designed to broaden their understanding of the various areas of molecular cell biology as well as developmental biology, neurobiology, immunology and the cell biology of disease.